Flora of Puma Canyon Ecological Reserve, southeast of Pinon Hills

Fig. 1. Left: Canbya candida, pygmy poppy, with a penny in the background for scale. Right: Opuntia basilaris var. brachyclada, short-joint beavertail. Pictures taken by Tom Chester on 14 April 2015. Click on the pictures for larger versions.

The 320 acre Puma Canyon Ecological Reserve is an achievement of the Transition Habitat Conservancy, which has protected a total of over 6000 acres in the Mojave Desert.

The checklist for the reserve is young and hence quite incomplete, compiled from only the following visits so far, from:

See the 14 and 26 April 2005 survey routes on an area map.

The union of the field surveys contains 155 taxa, including five taxa that are not yet confidently determined. An additional six taxa are given in the checklist below that were found in the Sheep Creek Wash area outside the Reserve parcel. Because only a small part of the entire 320 acres has been surveyed so far, and the surveyed areas have each only been covered once, it is certain that the list will grow with time as more surveys are done.

Michael Charters has made a beautiful photo gallery of 114 species from his visits in 2015. His gallery includes one species that we couldn't identify, and hence is not in the checklist given below. If you can identify it (it is at the bottom of page 9), please let us know!

In an attempt to make the following checklist more complete for future fieldwork, vouchers were searched at the Consortium of California Herbaria on 19 April 2015 in a larger area of similar habitat. First, a geographic coordinate search was made at the consortium using a latitude-longitude box larger than the area desired (note that this of course does not retrieve vouchers that have not been georeferenced). Then only the vouchers in the area south of SR138, north of the base of the San Gabriel Mountains, east of Puzzle Canyon, and west of the hills immediately west of SR2 were selected. Fig. 2 shows the locations of the accepted vouchers, along with an approximate rendition of the areas surveyed on 14 and 26 April 2015.

Fig. 2. An area map showing the number of vouchers at various locations (colored circles with numbers inside, from the clustering algorithm used by the Consortium of California Herbaria), with yellow circles showing locations with more than 10 vouchers and blue circles showing locations with fewer vouchers. The approximate routes surveyed on 14 and 26 April 2015 were drawn in by hand (heavy blue line; see more precise survey routes on the topo map). Note that no georeferenced vouchers have ever been collected along the route surveyed. The Angeles Crest Highway (SR2) is the white highway line at extreme lower left and at bottom right. The town of Wrightwood is just of the bottom edge of the map, with Cajon Pass just off the lower right edge of the map.

One species, Lessingia germanorum, was rejected from the voucher list as being probably misdetermined, since it is an extremely rare species found only at two small locations in San Francisco and San Mateo County.

The vouchers of another species, Mimulus bigelovii, are possibly actually of M. fremontii; see the discussion of the vouchers of Mimulus bigelovii and M. fremontii. See that page for photographs of the M. fremontii from the Puma Reserve, and a discussion of how to distinguish it from M. bigelovii.

A total of 152 taxa were found in the 375 accepted vouchers.

The vouchers are almost entirely the work of one person: Dick Swinney. He collected 312 out of the 375 vouchers, an amazing 83% of the total. Without Dick's work, very few additional species would have been added to the following checklist from the voucher search.

The total checklist below contains 223 taxa, 155 definitely known from the Reserve, with some, but not all, of the other 68 taxa expected to be found in the Reserve. It is also likely that future surveys of the reserve will find species not present in the voucher list.

The total checklist is very unusual in that there are no lycophytes or ferns!

The last three columns in the checklist below are as follows:

The species with uncertain determinations in the field have a qualifier in these columns, ~ for our best-guess determination and sp if the species is completely uncertain, with one vouchered Nemacladus species name just a place-holder for the observation of the genus in that case.

Version for printing, without lines and other text on this page: html (6 pages) or pdf Clickbook booklet (2 double-sided pages). (See printing instructions for an explanation of these options)

See Information about the order in which the species are presented, and the links from the Scientific Name and Common Name.

See also the beautiful photo gallery of 114 species from Michael Charters.

#FamilScientific Name(*)Common Name#V#Plants
1CupreJuniperus californicaCalifornia juniper45099
2CupreJuniperus osteospermaUtah juniper1 PM
3EphedEphedra nevadensisNevada ephedra 9930
4PinacPinus monophyllapinyon pine 2099
5ApiacLomatium mohavenseMojave lomatium3WF8
6ApiacLomatium nevadense var. parishiiParish's lomatium  10
7ApiacTauschia parishiiParish's tauschia5 10
8ApocyAsclepias californicaCalifornia milkweed  1
9ApocyAsclepias erosadesert milkweed  PM
10ApocyAsclepias vestitawoolly milkweed154
11AsterAcamptopappus sphaerocephalus var. hirtellushairy goldenhead2 2
12AsterAmbrosia acanthicarpabur-ragweed 3040
13AsterAmbrosia salsola var. salsolacheesebush4110
14AsterAnisocoma acaulisscale-bud1 15
15AsterArtemisia dracunculuswild tarragon  PM
16AsterArtemisia tridentatabig sagebrush 1099
17AsterBaccharis salicifolia ssp. salicifoliamule fat1  
18AsterBaileya pleniradiatawoolly marigold2  
19AsterCalycoseris parryiyellow tackstem21PM
20AsterChaenactis fremontiiFremont pincushion1  
21AsterChaenactis stevioidesdesert pincushion29999
22AsterChaenactis xantianaXantus' chaenactis2 PM
23AsterCirsium occidentale var. californicumCalifornia thistle  PM
24AsterEncelia actoniActon encelia 2099
25AsterEricameria cooperi var. cooperiCooper's goldenbush2  
26AsterEricameria linearifolianarrowleaf goldenbush31099
27AsterEricameria nauseosa var. ceruminosasharp-bracted rabbitbrush1  
28AsterEricameria nauseosa var. hololeucaghostly rabbitbrush 305
29AsterErigeron foliosus var. foliosusleafy daisy12050
30AsterEriophyllum confertiflorum var. confertiflorumgolden yarrow  10
31AsterEriophyllum pringleiPringle's woolly sunflower39999
32AsterEriophyllum wallacei var. wallaceiWallace's woolly daisy29999
33AsterGlyptopleura marginatacarved-seed1  
34AsterGrindelia squarrosa var. serrulata*curlycup gumweed1  
35AsterGutierrezia microcephalasticky snakeweed351
36AsterLasthenia gracilisgoldfields231
37AsterLayia glandulosawhite tidy-tips59999
38AsterLepidospartum latisquamumNevada broomsage 36
39AsterLepidospartum squamatumscale-broom 99 
40AsterLeptosyne bigeloviiBigelow's coreopsis4530
41AsterLessingia glandulifera var. glanduliferasticky lessingia 4099
42AsterMalacothrix glabratadesert dandelion332
43AsterSenecio flaccidus var. douglasiibush senecio2  
44AsterStephanomeria exigua ssp. coronariaslender wreathplant  99
45AsterStephanomeria exigua ssp. exiguaslender wreathplant2~3 
46AsterStephanomeria parryiParry's stephanomeria3 ~99
47AsterStephanomeria pauciflorawire-lettuce15PM
48AsterSyntrichopappus fremontiiFremont's syntrichopappus1  
49AsterTetradymia axillaris var. longispinacatclaw horsebrush2550
50AsterUropappus lindleyisilver puffs21010
51BoragAmsinckia tessellata var. tessellatabristly fiddleneck23099
52BoragCryptantha barbigera var. barbigerabearded cryptantha1  
53BoragCryptantha circumscissa var. circumscissacushion cryptantha130 
54BoragCryptantha decipiensgravel cryptantha 55
55BoragCryptantha micrantha var. micranthared-root cryptantha4  
56BoragCryptantha mohavensisMojave cryptantha1  
57BoragCryptantha muricataprickly cryptantha1  
58EudicCryptantha nevadensis var. rigidarigid cryptantha15PM
59BoragCryptantha oxygonasharp-nut cryptantha3 PM
60BoragCryptantha pterocarya var. pterocaryawing-nut cryptantha499 
61BoragCryptantha pterocarya var. purpusiiPurpus' wing-nut cryptantha 1099
62BoragCryptantha similiscushion cryptantha39999
63BoragEmmenanthe penduliflora var. penduliflorawhispering bells 401
64BoragEriodictyon trichocalyx var. trichocalyxsmoothleaf yerba santa 9930
65BoragNama californicaCalifornia fiddleleaf1  
66BoragNama demissa var. demissapurple mat  20
67BoragNama depressanarrow-leaved nama  PM
68BoragNemophila menziesii var. menziesiibaby blue eyes2 2
69BoragPectocarya penicillatacombseed4130
70BoragPectocarya setosamoth combseed59999
71BoragPhacelia cicutaria var. hispidacaterpillar phacelia  PM
72BoragPhacelia curvipescurve-stem phacelia311
73BoragPhacelia distanscommon phacelia23050
74BoragPhacelia fremontiiFremont's phacelia3550
75BrassBoechera pulchrabeautiful rock-cress65PM
76BrassBoechera xylopodaslender & beautiful rock-cress 1 
77BrassCaulanthus cooperiCooper's jewel-flower2  
78BrassCaulanthus lasiophyllusCalifornia mustard29999
79BrassDescurainia pinnatawestern tansy-mustard23010
80BrassDescurainia sophia*herb sophia  2
81BrassErysimum capitatum var. capitatumwestern wallflower2  
82BrassHirschfeldia incana*shortpod mustard 20 
83BrassLepidium fremontiidesert pepper-grass540 
84BrassSisymbrium altissimum*tumble-mustard19950
85BrassSisymbrium orientale*Oriental mustard N1
86BrassThysanocarpus laciniatuslace-pod322
87CactaCylindropuntia echinocarpasilver cholla 550
88CactaOpuntia basilaris var. brachycladashort-joint beavertail 111
89CampaNemacladus longiflorusthread plant1  
90CampaNemacladus ramosissimusNuttall's nemacladus1  
91CampaNemacladus sigmoideussmall-flowered nemacladus3 spPM
92CapriLonicera interruptachaparral honeysuckle2  
93CaryoEremogone macradeniadesert sandwort1 3
94CaryoSilene verecundawhite catch-fly1  
95ChenoChenopodium berlandieripit-seed goosefoot ~N 
96ChenoChenopodium californicumCalifornia goosefoot39999
97ChenoChenopodium fremontiiFremont's goosefoot N 
98ChenoGrayia spinosahop-sage N5
99ChenoKrascheninnikovia lanatawinter fat2 1
100ChenoSalsola tragus*Russian thistle 206
101ConvoCuscuta californica var. californicaCalifornia dodder1 ~99
102EricaArctostaphylos glaucabigberry manzanita  5
103EuphoChamaesyce albomarginatarattlesnake weed3WF5
104FabacAcmispon americanus var. americanusSpanish clover1  
105FabacAcmispon procumbens var. procumbenssilky California broom 120
106FabacAstragalus didymocarpusdwarf white milk-vetch  PM
107FabacAstragalus douglasiiJacumba milk-vetch3  
108FabacAstragalus lentiginosus var. variabilisfreckled milk-vetch61 
109FabacAstragalus pachypus var. pachypusthick-pod milk-vetch1  
110FabacLupinus concinnusbajada lupine N 
111FabacLupinus excubitus var. austromontanusmountain grape-soda lupine421
112FabacLupinus odoratusMojave lupine N 
113FabacMelilotus albus*white sweetclover 20 
114FagacQuercus john-tuckeriTucker's oak1 2
115GentiFrasera neglectapine green-gentian1  
116GeranErodium cicutarium*redstem filaree N20
117LamiaMonardella exilisMojave monardella  10
118LamiaSalvia columbariaechia11020
119LamiaSalvia dorrii var. dorriiblue sage2  
120LamiaSalvia dorrii var. pilosahairy sage 9999
121LamiaScutellaria mexicanaMexican bladder sage2302
122LoasaMentzelia albicauliswhite-stemmed blazing star29999
123LoasaMentzelia laevicaulisgiant blazing star 1 
124LoasaMentzelia veatchianaVeatch's blazing star3 20
125MalvaFremontodendron californicumflannelbush 520
126MalvaSphaeralcea ambigua var. ambiguaapricot mallow2  
127MontiCalyptridium monandrumsand cress5110
128MontiClaytonia parvifloranarrow-leaved miner's lettuce2  
129MontiClaytonia parviflora ssp. parvifloranarrow-leaved miner's lettuce  20
130MontiClaytonia perfoliataminer's lettuce  PM
131NyctaAbronia pogonanthaMojave sand verbena1  
132NyctaMirabilis laevis var. villosaBigelow's desert four-o'clock25099
133OnagrCamissonia campestris ssp. campestrisfield suncup39999
134OnagrCamissoniopsis bistortasouthern suncup  PM
135OnagrCamissoniopsis confusaSan Bernardino suncup1  
136OnagrCamissoniopsis pallida ssp. pallidapale suncup51 
137OnagrEremothera boothii ssp. decorticansBooth's evening primrose420 
138OnagrOenothera californicaCalifornia evening-primrose1  
139OrobaCastilleja plagiotomaMojave Indian paintbrush223
140OrobaOrobanche valida ssp. validaRock Creek broomrape  ~PM
141PapavArgemone munitaprickly poppy 1 
142PapavCanbya candidapygmy poppy  8
143PapavEschscholzia minutiflora ssp. covilleiCoville's poppy N 
144PapavEschscholzia minutiflora ssp. minutiflorasmall-flowered poppy 51
145PhrymMimulus androsaceusandrosace monkeyflower2  
146PhrymMimulus bigeloviiBigelow's monkeyflower3  
147PhrymMimulus fremontii var. fremontiiFremont's monkeyflower 5099
148PhrymMimulus johnstoniiJohnston's monkeyflower  50
149PhrymMimulus rubelluslittle redstem monkeyflower1  
150PlantAntirrhinum coulterianumCoulter's snapdragon 3PM
151PlantPenstemon centranthifoliusscarlet bugler114
152PolemEriastrum densifolium ssp. austromontanumsouthern mountain woolly-star  20
153PolemEriastrum eremicum ssp. eremicumdesert woolly-star 1050
154PolemEriastrum sapphirinumsapphire woolly-star6 20
155PolemGilia canashowy gilia33099
156PolemGilia minorlittle gilia1  
157PolemGilia modocensisModoc gilia2110
158PolemGilia ochroleuca ssp. bizonatavolcanic gilia  99
159PolemGilia sinuatarosy gilia  1
160PolemLeptosiphon breviculusMojave linanthus59999
161PolemLinanthus bigeloviiBigelow's linanthus2 1
162PolemLinanthus parryaesand-blossoms2 2
163PolemLoeseliastrum matthewsiidesert calico62040
164PolemSaltugilia splendens ssp. splendenssplendid gilia1 4
165PolygCentrostegia thurberired triangles34099
166PolygChorizanthe brevicornu var. brevicornubrittle spineflower1  
167PolygChorizanthe watsoniiWatson's spineflower2 10
168PolygEriogonum baileyi var. baileyiBailey's buckwheat2  
169PolygEriogonum brachyanthumshort-flowered buckwheat3  
170PolygEriogonum davidsoniiDavidson's buckwheat39999
171PolygEriogonum fasciculatum var. polifoliumCalifornia buckwheat59999
172PolygEriogonum heermannii var. heermanniiHeermann's buckwheat523
173PolygEriogonum maculatumspotted buckwheat5  
174PolygEriogonum mohavensewestern Mojave buckwheat1  
175PolygEriogonum plumatellayucca buckwheat221
176PolygEriogonum pusillumyellow-turbans59950
177PolygEriogonum roseumwand buckwheat1  
178PolygEriogonum saxatilerock buckwheat2  
179RanunDelphinium parishii ssp. parishiiParish's larkspur73099
180RhamnCeanothus vestitusMojave ceanothus3 1
181RhamnFrangula californica ssp. cuspidataSierra hoary coffeeberry2N 
182RosacPrunus fasciculata var. fasciculatadesert almond19999
183RosacPurshia tridentata var. glandulosadesert bitterbrush59910
184RubiaGalium angustifolium ssp. angustifoliumnarrowleaf bedstraw3105
185RubiaGalium johnstoniiJohnston's bedstraw1  
186SalicPopulus fremontii ssp. fremontiiFremont cottonwood21 
187SalicSalix laevigatared willow1  
188SalicSalix lasiolepisarroyo willow6  
189SolanLycium andersoniiAnderson's boxthorn1  
190SolanLycium cooperiCooper's boxthorn2 2
191SolanSolanum rostratum*buffalo berry1  
192ViolaViola purpurea ssp. mohavensisMojave goosefoot violet2  
193ViscaPhoradendron bolleanumdense mistletoe32010
194AgavaHesperoyucca whippleichaparral yucca 2050
195AgavaYucca brevifoliajoshua tree29999
196AlliaAllium fimbriatum var. fimbriatumfringed onion3WF99
197CyperCarex almasturdy sedge3  
198CyperSchoenoplectus acutus var. occidentaliscommon tule1  
199JuncaJuncus balticus ssp. aterwire rush4  
200JuncaJuncus nodosusknotted rush1  
201LiliaCalochortus kennedyi var. kennedyired mariposa 130
202PoaceArundo donax*giant reed1  
203PoaceBromus carinatusCalifornia brome2  
204PoaceBromus madritensis ssp. rubens*red brome 9950
205PoaceBromus tectorum*downy brome19999
206PoaceElymus elymoides var. elymoidessquirreltail1  
207PoaceElymus triticoidescreeping wild rye2  
208PoaceFestuca microstachyssmall fescue1  
209PoaceFestuca octoflorasix-weeks fescue3  
210PoaceKoeleria macranthaJune grass  20
211PoacePoa bulbosa ssp. vivipara*bulbous blue grass  PM
212PoacePoa pratensis*Kentucky blue grass2  
213PoacePoa secundaone-sided blue grass1  
214PoacePoa secunda ssp. secundaone-sided bluegrass 9999
215PoaceSchismus arabicus*Mediterranean grass  PM
216PoaceSchismus barbatus*Mediterranean schismus3  
217PoaceStipa hymenoidesIndian ricegrass  1
218PoaceStipa parishii var. parishiiParish's needlegrass1  
219PoaceStipa speciosadesert needlegrass 2030
220PoaceTriticum aestivum*common wheat  1
221ThemiDichelostemma capitatum ssp. capitatumblue dicks 110
222ThemiMuilla coronatacrowned muilla1  
223ThemiMuilla maritimamuilla  5

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Copyright © 2015 by Tom Chester, Michael Charters, RT Hawke, Pam MacKay, Tom Egan, Tim Thomas, Wendy Walker, Nancy Accola, Adrienne Ballway, Walt Fidler, Frank Harris, Shaun Hawke and Harry Spilman.
Permission is freely granted to reproduce any or all of this page as long as credit is given to us at this source:
Comments and feedback: Tom Chester
Last update: 4 June 2015