Plant Guide to Guy Fleming Trail, Torrey Pines State Reserve This is a working list, about which I make no guarantees at all until I officially release it. Use at your own risk!
Introduction and Explanation of Plant Trail Guides
Introduction
Highlights of This Trail
Plant Communities and Floristics
Fieldwork Dates and Summary of List Changes With Time
Botanical Trip Reports
The Plant Guide
Comments On Specific Species
Introduction See Introduction to Torrey Pines Trails in the Beach Trail Plant Guide.
Highlights of This Trail The botanical highlights of this trail are:
Plant Communities and Floristics This section is not updated every time the guide is updated. Last update for this section: 27 February 2004. (Only the histogram in the Number of Unique Taxa On This Trail is always updated each time the guide is updated.)
See Plant Communities and Floristics for the Beach Trail.
The Trail plant list contains 96 taxa, of which 90 have so far been identified, in 0.87 unique miles of trail. One additional taxon has not been identified to species, but it is a non-native iceplant. Of these 91 taxa, 14 (15%) are non-native.
I have plotted these numbers against the other trails in my database. The plots show:
- For its elevation, the percentage of native taxa is surprisingly high compared to the other trails in our database. This is most likely due to the relative pristineness of this area, but the incompleteness of this trail guide so far could also be a factor. The subnormal rainfall in the years I've surveyed this trail might not have germinated all the non-native annuals.
- The number of native taxa is about that expected from our other trails.
Summary statistics:
By Number
Lifeform Native Non-native Total Taxa Trees 1 0 1 Shrubs 29 2 31 Pteridophytes 2 0 2 Perennial Herbs 25 2 27 Perennial Grasses 4 0 4 Annual Grasses 0 5 5 Annual Herbs 16 5 21 Total 77 14 91 By Percent
Lifeform Native Non-native Total Taxa Trees 1% 0% 1% Shrubs 38% 14% 34% Pteridophytes 3% 0% 2% Perennial Herbs 32% 14% 30% Perennial Grasses 5% 0% 4% Annual Grasses 0% 36% 5% Annual Herbs 21% 36% 23% Total 100% 100% 100% Number of Unique Taxa On This Trail
The following histogram gives the number of trails in my database that contain each taxon on this trail. I had 73 trails in my database when this histogram was made; 2 of those trails, including this one, are at Torrey Pines. A number of "1" means the taxon has only been found on this trail among the trails in my database; numbers of "2" or smaller may indicate taxa found only at Torrey Pines.
Number of Trails
Containing A TaxonNumber Of Taxa
On This Trail% of Taxa
On This Trail1 2 2% 2 15 13% 3 4 3% 4 5 4% 5 7 6% 1-5 33 29% 6-10 16 14% 11-15 18 16% 16-20 11 10% 21-25 11 10% 26-30 5 4% 31-35 3 3% 36-40 8 7% 41-45 6 5% 46-50 4 3% Total Taxa 115 100% I found 6 additional species not in the above table, since they have not been identified yet. The unidentified ones are marked with ? or sp in the id? column in the guide, and have no entries in the #all column.
The taxa that are truly unique, or almost unique, to this trail or area, out of the 73 trails in my database on 7 February 2004, are:
The following taxon is listed as being found only on this trail, but was planted here:
#all Latin Name Common Name Comment 1 Agave americana *American agave A handful of specimens have been planted along the main road
Fieldwork Dates and Summary of List Changes With Time The following table gives the dates the trail was walked and taxa recorded. After each visit, the table gives the total number of taxa on the list and the breakdown of the taxa without positive identification. See Explanation of Plant Trail Guides to understand the symbols below.
Visit Date Visit # # taxa # "?" # "sp" # "~" # "ssp" 6/18/2003 1 81 4 7 3 2 11/29/2003 2 87 3 7 1 1 2/5/2004 3 96 2 6 3 1 3/18/2004 4 111 2 4 4 0 1/13/2005 5 121 2 4 9 0 I thank James Dillane for help with the 11/29/03 fieldwork, especially for correcting one error in the guide and for supplying the names of other species found on this trail that were not visible in June or November.
I thank Jane Strong for help with the 2/5/04 fieldwork, for helping to find and identify new species on the trail, as well as verifying the species in the guide.
I thank a docent at Torrey Pines, who wishes to remain anonymous, for her help with the 1/13/05 fieldwork for the first 0.2 miles.
Botanical Trip Reports 5 February 2004
18 February 2004
The Plant Guide Version for printing, without lines and other text on this page (6 pages)
Mile S # id? Common Name Latin Name #here #all 0.00 Beginning of trail, at the first Torrey Pine; elevation ~245 feet (75 m). 0.00 b 1 Torrey pine Pinus torreyana 20 / 9 2 0.00 r 2 *American agave Agave americana 5 / 1 1 0.00 l 3 *purple false-brome Brachypodium distachyon / 5 0.00 l 4 wild-cucumber Marah macrocarpus var. macrocarpus 5 / 5 40 0.00 l 5 narrowleaf bedstraw Galium angustifolium ssp. angustifolium 20 / 4 41 0.00 l 6 *smooth cat's ear Hypochaeris glabra / 27 0.00 b 7 coast prickly-pear Opuntia littoralis 2 / 2 7 0.00 r 8 climbing bedstraw Galium nuttallii ssp. nuttallii 10 / 9 28 0.00 r 9 deerweed Lotus scoparius var. scoparius 30 / 9 22 0.00 l 10 black sage Salvia mellifera 10 / 9 37 0.00 l 11 purple snapdragon Antirrhinum nuttallianum ssp. nuttallianum 99 / 9 12 0.00 l 12 California-aster Lessingia filaginifolia var. filaginifolia+ 30 / 9 50 0.00 l 13 saw-toothed goldenbush Hazardia squarrosa var. grindelioides 20 / 9 32 0.00 b 14 California buckwheat Eriogonum fasciculatum var. foliolosum 30 / 9 46 0.00 r Sign: "Stay on designated trail. Reserve hours 7 a.m. to sunset." 0.00 r 15 California sagebrush Artemisia californica 10 / 5 42 0.00 l 16 giant needlegrass Achnatherum coronatum 20 / 9 21 0.00 l 17 lanceleaf dudleya Dudleya lanceolata 50 / 9 19 0.00 l 18 ? grass similar to poa secunda ? 10 / 1 0.00 l 19 pygmy-weed Crassula connata+ 50 / 3 14 0.00 r 20 foothill needlegrass Nassella lepida 20 / 9 18 0.00 r 21 San Diego wreathplant Stephanomeria diegensis 40 / 9 18 0.00 b 22 bush monkeyflower Mimulus aurantiacus 1 / 1 39 0.01 l 23 toyon Heteromeles arbutifolia 2 / 2 42 0.01 l 24 *Mediterranean pygmy-weed Crassula tillaea+ 30 / 1 5 0.01 l 25 *Australian brass-buttons Cotula australis 50 / 3 14 0.01 b 26 milk maids Cardamine californica+ 40 / 9 8 0.01 r 27 *redstem filaree Erodium cicutarium 99 / 9 48 0.01 b 28 lemonade berry Rhus integrifolia 40 / 9 12 0.01 l 29 southern honeysuckle Lonicera subspicata var. denudata 1 / 1 38 0.01 r 30 popcorn flower Cryptantha intermedia 99 / 9 22 0.01 r (laurel sumac, Malosma laurina) 0.01 l 31 bushrue Cneoridium dumosum 40 / 9 3 0.01 r 32 *red brome Bromus madritensis ssp. rubens 20 / 2 44 0.01 r 33 ? annual everlasting? Gnaphalium stramineum? 20 / 3 0.01 r Check for California chicory, Rafinesquia californica 0.02 r 34 wartystem ceanothus Ceanothus verrucosus 50 / 9 3 0.02 r 35 San Diego morning-glory Calystegia macrostegia ssp. tenuifolia 10 / 5 17 0.02 l 36 sea dahlia Coreopsis maritima 50 / 9 4 0.02 l 37 redberry Rhamnus crocea 20 / 5 15 0.02 l Bench. 0.02 l Display board. 0.02 Y-jct. Go right. 0.02 l 38 sp suncup Camissonia sp. 99 / 9 0.02 b 39 Mohave yucca Yucca schidigera 20 / 9 6 0.02 l 40 threadstem Pterostegia drymarioides 50 / 4 20 0.03 l 41 San Diego barrel cactus Ferocactus viridescens 3 / 3 2 0.03 r (western prickly pear, Opuntia occidentalis) 0.03 r 42 (can't read my writing....) 0.03 r 43 ~ everlasting nest-straw Stylocline gnaphaloides 99 / 9 7 0.03 r 44 ~ *Mediterranean schismus Schismus barbatus 20 / 2 16 0.03 r 45 chamise Adenostoma fasciculatum 10 / 9 35 0.04 r 46 shiny lomatium Lomatium lucidum 10 / 3 6 0.04 Begin fence posts on right side of trail. 0.04 r (Parish's purple nightshade, Solanum parishii) 0.05 r 47 mission manzanita Xylococcus bicolor 3 / 3 12 0.06 l 48 *annual veldt grass Ehrharta longiflora+ 99 / 9 2 0.06 l 49 leafy daisy Erigeron foliosus var. foliosus 5 / 3 34 0.06 l 50 *common groundsel Senecio vulgaris 10 / 2 14 0.06 l Sign: "Toyon, Heteromeles arbutifolia". 0.06 l 51 yellow mariposa lily Calochortus weedii var. weedii 30 / 9 7 0.06 l 52 ~ seashore bentgrass Agrostis pallens 99 / 4 5 0.06 l 53 southern Indian pink Silene laciniata ssp. major / 15 0.06 l 54 California polypody Polypodium californicum 99 / 5 15 0.06 l 55 ladies fingers Dudleya edulis 50 / 9 7 0.06 l 56 small-flowered soap plant Chlorogalum parviflorum 10 / 2 5 0.06 Four steps up. 0.07 b 57 golden yarrow Eriophyllum confertiflorum var. confertiflorum 20 / 9 50 0.07 l (coast scrub oak, Quercus dumosa) 0.07 Four steps down. 0.08 l 58 woolly Indian paintbrush Castilleja foliolosa 3 / 2 4 0.08 b 59 blue-eyed grass Sisyrinchium bellum 10 / 1 18 0.08 l 60 shooting star Dodecatheon clevelandii ssp. clevelandii 10 / 1 10 0.08 l Sign: "chamise, Adenostoma fasciculatum" 0.08 r 61 Orcutt's yellow pincushion Chaenactis glabriuscula var. orcuttiana+ / 2 0.08 b 62 coastal goldenbush Isocoma menziesii+ 3 / 2 13 0.09 r 63 bicolored everlasting Gnaphalium bicolor 3 / 2 27 0.09 r 64 ~ *narrowleaf filago Filago gallica / 41 0.10 r Sign: "Birdbath is filled several times per week." 0.10 r 65 ~ pink sand verbena Abronia umbellata ssp. umbellata+ 30 / 9 2 0.10 l Drinking fountain. 0.11 l Low bench. 0.11 r 66 *creeping Australian saltbush Atriplex semibaccata 3 / 2 10 0.13 l 67 Parish's purple nightshade Solanum parishii 20 / 9 16 0.13 l 68 blue dicks Dichelostemma capitatum ssp. capitatum 20 / 2 40 0.13 b 69 western dichondra Dichondra occidentalis 20 / 2 2 0.14 r (mission cactus, spineless form, Opuntia ficus-indica) 0.15 l 70 California poppy Eschscholzia californica 20 / 7 24 0.15 l 71 sandmat Cardionema ramosissimum 20 / 9 2 0.15 l 72 silverback fern Pentagramma triangularis ssp. viscosa 10 / 1 4 0.15 l (wide-throated yellow monkeyflower, Mimulus brevipes) 0.17 l (white sage, Salvia apiana) 0.17 r 73 south coast morning-glory Calystegia macrostegia ssp. intermedia 5 / 5 15 0.18 r 74 bristly bird's beak Cordylanthus rigidus ssp. setigerus 30 / 9 25 0.18 l 75 ~ California filago Filago californica / 15 0.18 l Sign: "black sage, Salvia mellifera". 0.19 b 76 ~ *scarlet pimpernel Anagallis arvensis 20 / 2 23 0.20 l 77 miner's lettuce Claytonia perfoliata ssp. perfoliata 5 / 2 21 0.21 l 78 wild celery Apiastrum angustifolium 20 / 2 10 0.22 l 79 *rattail fescue Vulpia myuros var. myuros 5 / 1 11 0.22 l 80 sp western wallflower? Erysimum capitatum ssp. capitatum?+ 10 / 3 0.22 l 81 California plantain Plantago erecta 50 / 3 10 0.23 l 82 narrow-leaved miner's lettuce Claytonia parviflora ssp. parviflora / 15 0.27 l 83 ~ coast-range melic Melica imperfecta / 37 0.28 l 84 common bedstraw Galium aparine 20 / 2 37 0.28 l 85 *common chickweed Stellaria media 20 / 1 21 0.28 l 86 San Diego fiesta flower Pholistoma racemosum 50 / 3 2 0.28 l Sign: "scrub oak, Quercus dumosa". 0.28 b 87 coast scrub oak Quercus dumosa 2 / 2 2 0.28 l 88 graceful bedstraw Galium porrigens var. porrigens 5 / 1 4 0.28 r 89 *mission cactus (spiny form) Opuntia ficus-indica 1 / 1 5 0.29 b 90 saltgrass Distichlis spicata 50 / 4 14 0.30 Low bridge under a tree branch 0.30 r 91 sp annual goosefoot Chenopodium sp. 5 / 1 0.31 l 92 three spot Osmadenia tenella 30 / 2 7 0.32 l 93 dwarf lupine Lupinus bicolor 1 / 1 26 0.32 r Jct. ocean overlook trail. Go right and take it. Due to the width of the trail, look only the plants on the right, both going to the overlook and coming back. 0.33 r 94 California croton Croton californicus 10 / 5 6 0.33 Turn around at end of overlook trail; elevation ~230 feet (70 m). 0.34 r 95 California encelia Encelia californica 20 / 5 12 0.35 r 96 *Crete weed Hedypnois cretica / 16 0.35 r 97 shrubby beach-primrose Camissonia cheiranthifolia ssp. suffruticosa 20 / 5 2 0.35 r (johnny jump-up, Viola pedunculata) 0.35 r (common tidy-tips, Layia platyglossa) 0.35 r 98 prostrate spineflower Chorizanthe procumbens / 3 0.35 r 99 California spineflower Mucronea californica 5 / 2 1 0.35 r 100 ~ meally white pincushion Chaenactis artemisiifolia 1 / 1 13 0.36 Jct. main trail, go right. 0.37 Sign: "California buckwheat, Eriogonum fasciculatum". 0.37 l 101 strigose lotus Lotus strigosus 1 / 1 25 0.38 Trail curves left 90° 0.39 r 102 Robinson's pepper-grass Lepidium virginicum var. robinsonii 20 / 3 5 0.39 r (coast cholla, Opuntia prolifera) 0.39 r 103 climbing milkweed Sarcostemma cynanchoides ssp. hartwegii 50 / 5 2 0.39 b 104 bladderpod Isomeris arborea 20 / 5 7 0.39 l 105 *Russian thistle Salsola tragus 20 / 3 25 0.40 l 106 collar lupine Lupinus truncatus 2 / 1 20 0.40 l 107 *ripgut brome Bromus diandrus / 44 0.40 l 108 ground pink Linanthus dianthiflorus 20 / 2 8 0.40 Trail curves right 90° 0.41 l 109 western prickly-pear Opuntia occidentalis 2 / 2 2 0.41 l 110 *slender wild oats Avena barbata / 25 0.42 r Sign: "Bush sunflower, Encelia californica". 0.42 r (blue larkspur, Delphinium parryi ssp. maritimum) 0.43 l Bench without a back. 0.44 r Sign: "Torrey Pine, Pinus torreyana". 0.44 r 111 south coast branching phacelia Phacelia ramosissima var. austrolitoralis 10 / 3 3 0.44 Trail curves left. 0.44 l 112 California groundsel Senecio californicus 20 / 2 2 0.46 r 113 *crystalline ice plant Mesembryanthemum crystallinum 50 / 3 5 0.46 r Sign: "Mojave Yucca, Yucca schidigera". 0.48 l Blank sign 0.50 l (Shaw's agave, Agave shawii) 0.50 l Bench without a back. 0.50 l Sign: "sea-cliff buckwheat, Eriogonum parvifolium". 0.50 l 114 sea-cliff buckwheat Eriogonum parvifolium 3 / 3 4 0.51 l 115 *sea fig Carpobrotus chilensis+ 1 / 1 2 0.57 l Bench without a back. 0.57 Y-jct. Go right. 0.61 Observation Point; elevation ~220 feet (65 m). Turn around. 0.65 Jct. main trail, go right. 0.65 l Sign: "Dead pines. The tree skeletons are all that remain from severe drought and bark beetle outbreak in the mid 1960s." 0.66 l 116 laurel sumac Malosma laurina 1 / 1 38 0.67 l Sign: "Mojave Yucca, Yucca schidigera". 0.68 l 117 sp California dodder Cuscuta californica var. californica 2 / 2 14 0.71 l Sign: "redberry, Rhamnus crocea". 0.72 l 118 Shaw's agave Agave shawii 3 / 1 2 0.72 l Bench without a back 0.74 Trail turns left 90° 0.74 r 119 California four o'clock Mirabilis californica 3 / 1 18 0.74 r 120 chaparral yucca Yucca whipplei 1 / 1 27 0.76 Trail turns left 90° 0.77 Trail curves right 90° 0.78 l Bench without a back. 0.80 "Low bridge" under a dead pine. 0.83 Trail makes a long curve left. 0.85 End loop trail. 0.87 Trailhead. Present on trail, but unknown location: 121 goldfields Lasthenia californica / 7
Comments On Specific Species Lessingia filaginifolia var. filaginifolia. The Jepson Manual combined two taxa that are found at Torrey Pines, Corethrogyne filaginifolia varieties linifolia and virgata. The editors at the Jepson Online Interchange have restored the Corethrogyne treatment, but have still dispensed with the varieties.
Because the var. linifolia is strikingly different from the usual var. seen in Southern California, var. virgata, I attempted to note which of the varieties is found on the nearby Beach Trail. However, when I attempted to use the key in Munz to key a plant to variety, it turned out to have characteristics of both varieties. Perhaps this is why the Jepson Manual dispensed with the varieties!
Crassula connata and C. tillaea. In the same location, Jane Strong found C. connata on 2/5/04 and I found C. tillaea 6 weeks later on 3/18/04! I verified C. connata on 1/13/05 and will try to verify C. tillaea in the future.
Cardamine californica. I studied the plants on this trail intensively on 1/13/05, and they do not fit either subspecies. Using the characteristics in the Jepson Manual, I found roughly as many plants that could be called var. californica as could be called var. integrifolia, with most plants not fitting either variety.
At the first location on this trail, I counted 23 oblong, ~entire to 2-toothed leaflets (var. integrifolia); and 26 ovate, mostly-toothed leaflets var. californica. In many cases, an individual plant had both kinds of leaves on it. Typically, the lowermost 1-2 cauline leaves would be ovate and toothed; the uppermost 1-2 cauline leaves would be oblong and ~entire, usually only with a terminal tooth. The leaves were intermediate between thin and thick. If one felt the blade as a whole, they were thickish, primarily due to the midvein. If one felt only the parts of the blade away from the midvein, they were thinnish. I surveyed leaves through the trail, finding the same results.
Note that var. integrifolia in the Jepson Manual is much changed from what it was in Munz. Using the Munz key, all these plants are var. californica, as were all Southern California plants in Munz. In the JM, integrifolia is now supposedly in Southern California, and there are vouchers determined as var. integrifolia in San Diego County.
I'll pay attention to milkmaids in other areas of Southern California, to see if any differ from these plants.
Chaenactis glabriuscula var. orcuttiana. The identification is from plants at mile 0.48 in 2002.
Isocoma menziesii. In my experience, the subspecies are not separable in Southern California. See Comments on the Jepson Manual and A Flora of Southern California by Munz: Isocoma menziesii.
Abronia umbellata ssp. umbellata. The first occurrence on the trail was a baby plant on 1/13/05; a definite identification was at mile 0.32.
Erysimum capitatum ssp. capitatum?. These plants are definitely wallflowers. The identification is uncertain because E. ammophilum also is supposed to occur here. I haven't yet checked the immature fruit for the id.
Carpobrotus chilensis. This specimen was weeded out on 3/l8/04.
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Copyright © 2003-2005 by Tom Chester.
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Comments and feedback: Tom Chester
Updated 15 January 2005.