Flora of Southern California: Lycopods and Ferns
Table of Contents
Brief Introduction to this Checklist
Notes on Some Taxa
Brief Introduction to this Checklist
See Flora of Southern California: Master Checklist for a more complete Introduction to this page.
The plants in this list are found on pages 109 to 134 in the printed JM2, with additional species found only in the online JM2 manual. See the notes below.
Click on the Scientific Name to get the JM2 online entry for each taxon, which includes:
- the JM2 keys and descriptions, with a list of all species found in a given genus or family, not just those found in southern California (no illustrations are online yet);
- a geographic map of the JM2 distribution;
- a plot of voucher locations, along with a shaded geographic area that fits the JM2 geographic and elevational distributions and hence indicates the potential geographic range for each taxon;
- a plot of elevation vs. latitude for vouchers, which is often a very instructive graph showing the change in the elevation range for each taxon with latitude; and
- a histogram of voucher collection by month, which often indicates when a taxon was in an identifiable state, either in bloom or in fruit.
Some links to the JM2 online entry will return no JM2 online entry, since hybrids, and the taxa mentioned as judgment-reserved taxa, are not yet entered into the JM2 name index. Those links may work in the future.
Click on the Common Name to get the Calphotos pictures. Since some taxa have no photos at Calphotos, those links will return no pictures. Be aware that the names in Calphotos are not synonymized, which means that if the name has changed in the JM2, all the Calphotos may still be under the JM1 name, and will not be retrieved by this link.
The column with header #Obs gives the number of observations in my database of each taxon from plant trail guides and floras that I've done.
The column #Lists gives the number of checklists that I've digitized that contain each taxon.
See Flora of Southern California: Master Checklist for some of the biases in those numbers.
# Family Scientific Name (*)Common Name #Obs #Lists Lycopods 1 Isoetaceae Isoetes bolanderi Bolander's quillwort 0 1 2 Isoetaceae Isoetes howellii Howell's quillwort 0 7 3 Isoetaceae Isoetes howellii var. minima Howell's quillwort 0 0 4 Isoetaceae Isoetes nuttallii Nuttall's quillwort 0 4 5 Isoetaceae Isoetes orcuttii Orcutt's quillwort 1 9 6 Selaginellaceae Selaginella asprella bluish spike-moss 5 19 7 Selaginellaceae Selaginella bigelovii Bigelow's spike-moss 27 62 8 Selaginellaceae Selaginella cinerascens ashy spike-moss 3 15 9 Selaginellaceae Selaginella eremophila desert spike-moss 30 19 10 Selaginellaceae Selaginella leucobryoides Mojave spike-moss 0 0 11 Selaginellaceae Selaginella watsonii Watson's spike-moss 11 19 Ferns 12 Aspleniaceae Asplenium septentrionale northern spleenwort 1 1 13 Aspleniaceae Asplenium vespertinum western spleenwort 0 13 14 Azollaceae Azolla filiculoides mosquito fern 0 19 15 Azollaceae Azolla microphylla^ Mexican mosquito fern 0 0 16 Blechnaceae Woodwardia fimbriata giant chain fern 3 38 17 Dennstaedtiaceae Pteridium aquilinum var. pubescens bracken 66 54 18 Dryopteridaceae Cyrtomium falcatum *Japanese netvein hollyfern 0 7 19 Dryopteridaceae Dryopteris arguta California wood fern 17 49 20 Dryopteridaceae Dryopteris filix-mas male fern 0 1 21 Dryopteridaceae Polystichum californicum California sword fern 0 0 22 Dryopteridaceae Polystichum imbricans cliff sword fern 0 7 23 Dryopteridaceae Polystichum imbricans ssp. curtum cliff sword fern 6 19 24 Dryopteridaceae Polystichum imbricans ssp. imbricans rock sword fern 0 4 25 Dryopteridaceae Polystichum kruckebergii Kruckeberg's sword fern 0 1 26 Dryopteridaceae Polystichum munitum western sword fern 0 5 27 Dryopteridaceae Polystichum scopulinum mountain holly fern 4 8 28 Equisetaceae Equisetum arvense field horsetail 3 23 29 Equisetaceae Equisetum hyemale ssp. affine horsetail 14 29 30 Equisetaceae Equisetum laevigatum smooth scouring rush 8 38 31 Equisetaceae Equisetum telmateia ssp. braunii giant horsetail 0 17 32 Equisetaceae Equisetum Xferrissii Ferris' horsetail 0 1 33 Marsileaceae Marsilea vestita ssp. vestita clover fern 3 15 34 Marsileaceae Pilularia americana pillwort 0 7 35 Ophioglossaceae Botrychium crenulatum scalloped moonwort 0 4 36 Ophioglossaceae Botrychium simplex var. simplex little grapefern 0 2 37 Ophioglossaceae Ophioglossum californicum California adder's-tongue 1 6 38 Ophioglossaceae Psilotum nudum *skeleton fork fern 0 3 39 Polypodiaceae^ Polypodium californicum California polypody 19 51 40 Polypodiaceae Polypodium hesperium western polypody 11 15 41 Polypodiaceae Polypodium scouleri leather-leaf fern 0 0 42 Pteridaceae Adiantum aleuticum five-fingered fern 1 6 43 Pteridaceae Adiantum capillus-veneris maidenhair fern 6 35 44 Pteridaceae Adiantum jordanii California maidenhair fern 7 35 45 Pteridaceae Argyrochosma jonesii Jones' false cloak-fern 0 4 46 Pteridaceae Argyrochosma limitanea ssp. limitanea^ cloak fern 0 0 47 Pteridaceae Aspidotis californica California lace fern 10 27 48 Pteridaceae Aspidotis densa Indian's dream 0 2 49 Pteridaceae Astrolepis cochisensis scaly cloak fern 0 0 50 Pteridaceae Cheilanthes clevelandii scaly lipfern 5 18 51 Pteridaceae Cheilanthes cooperae Mrs. Cooper's lipfern 0 3 52 Pteridaceae Cheilanthes covillei beady lipfern 29 64 53 Pteridaceae Cheilanthes covillei X C. parryi' teensy lipfern 0 4 54 Pteridaceae Cheilanthes feei slender lipfern 0 1 55 Pteridaceae Cheilanthes intertexta coastal lip fern 0 0 56 Pteridaceae Cheilanthes newberryi cottony fern 5 26 57 Pteridaceae Cheilanthes parryi woolly lipfern 50 45 58 Pteridaceae Cheilanthes viscida sticky lipfern 8 25 59 Pteridaceae Cheilanthes wootonii Wooton's lace fern 1 0 60 Pteridaceae Cryptogramma acrostichoides American parsley fern 1 6 61 Pteridaceae Notholaena californica California cloak fern 18 30 62 Pteridaceae Notholaena californica ssp. californica California cloak fern 0 0 63 Pteridaceae Notholaena californica ssp. leucophylla California cloak fern 0 0 64 Pteridaceae Pellaea andromedifolia coffee fern 24 57 65 Pteridaceae Pellaea breweri Brewer's cliff-brake 0 1 66 Pteridaceae Pellaea mucronata bird's-foot fern 0 11 67 Pteridaceae Pellaea mucronata var. californica California cliff-brake 9 18 68 Pteridaceae Pellaea mucronata var. mucronata bird's-foot fern 56 65 69 Pteridaceae Pellaea truncata cliff brake 1 4 70 Pteridaceae Pentagramma triangularis goldback fern 1 11 71 Pteridaceae Pentagramma triangularis ssp. maxonii^ Maxon's silverback fern 4 14 72 Pteridaceae Pentagramma triangularis ssp. rebmanii^ Rebman's silverback fern 0 0 73 Pteridaceae Pentagramma triangularis ssp. semipallida pale goldback fern 0 0 74 Pteridaceae Pentagramma triangularis ssp. triangularis goldback fern 35 53 75 Pteridaceae Pentagramma triangularis ssp. viscosa silverback fern 8 11 76 Pteridaceae Pteris cretica *Cretan brake 0 0 77 Pteridaceae Pteris tremula *brake 0 2 78 Pteridaceae Pteris vittata *ladder brake 1 5 79 Salviniaceae Salvinia minima^ *water spangles 0 0 80 Salviniaceae Salvinia molesta^ *giant salvina 0 2 81 Salviniaceae Salvinia oblongifolia^ *floating fern 0 0 82 Thelypteridaceae Thelypteris puberula var. sonorensis Sonoran maiden fern 0 7 83 Woodsiaceae^ Athyrium filix-femina var. cyclosorum subarctic lady-fern 20 12 84 Woodsiaceae^ Cystopteris fragilis brittle bladder fern 42 33 85 Woodsiaceae^ Woodsia oregana Oregon cliff fern 1 6 86 Woodsiaceae^ Woodsia plummerae Plummer's woodsia 0 3 87 Woodsiaceae^ Woodsia scopulina cliff fern 0 1
Notes on Some Taxa
The taxa with notes are given in alphabetical order here.
Cheilanthes covillei X C. parryi. This is a rare F1 hybrid between the two species, and has been given the name C. xparishii, mentioned in the JM2 treatment for C. covillei.
Cheilanthes intertexta. Not given as a southern California taxa in JM1 or JM2, but there are now nine vouchers at the Consortium for southern California, from SnBr near Lake Arrowhead; from Andreas Canyon near Palm Springs; and from Cuyamaca Peak in the Cuyamaca / Laguna Mountains. One voucher was determined by Steve Boyd; another by Jon Rebman, both quite knowledgeable about fern species.
However, one of those vouchers was determined by Munz as C. intertexta in 1922, yet Munz did not place this species in southern California in his 1974 Flora of Southern California.
I personally know nothing about C. intertexta, but in the JM2 key it seems extremely close to C. clevelandii, a very common species in southern California:
5. 2-colored rhizome scales with well-defined, dark mid-stripe and light brown margins 1/2 to = width of mid-stripe; leaf gray-green, segments round to ± cordate ..... C. clevelandii
5'. 2-colored rhizome scales with well-defined, dark mid-stripe and light brown margins much narrower than mid-stripe; leaf dark green, segments ± round ..... C. intertexta
As a result, it wouldn't be terribly surprising if a very small number of C. clevelandii plants had characteristics closer to those in the key for C. intertexta.
It would be interesting to know if the JM2 author knew about these vouchers and rejected this determination, or if the JM2 author was unaware of these vouchers.
Pentagramma triangularis ssp. maxonii. This taxon is marked as having a change, but the change was not in its name. What has changed are voucher determinations. Many vouchers formerly determined as ssp. viscosa in southwest San Diego County have now been redetermined as ssp. maxonii.
Polypodium scouleri. Given as being in SCo, n ChI in the JM2, with this note: Pls called P.s. from n ChI are fertile or sterile hybrids involving P. californicum. SCo was added to its geographic range in JM2. It would be interesting to know if those plants are also hybrids.
Copyright © 2012 by Tom Chester
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Last update: 21 February 2012